The scientists introduced genes from two other flowers that allowed the mums to mimic the chemical process producing blue pigment. This might be applicable to other flowers, like roses and lilies.
What are some plants that are mostly or completely red (not
Christian mythology, Vegder's Blog
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True Blue Chrysanthemum Flowers Produced with Genetic Engineering
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23279834 - HortScience) (372) Water Requirements of High Bush
For a New World to Come: Experiments in Japanese Art and
Japanese scientists create a true blue chrysanthemum
The world's first blue chrysanthemums created in Japan •
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The breakthrough behind those 'pure blue' chrysanthemums was pure luck